
Ling Ling was 22 years and seven months old, equivalent to about 70 human years, the Ueno Zoo said. It said he was the fifth-oldest known male panda in the world.
Keepers and visitors mourned the panda, which was the zoo's most popular attraction for more than 15 years. Public broadcaster NHK showed many visitors writing condolence messages, with some brushing away tears.
"He always turned around when I called 'Ling Ling.' This morning he didn't. I rushed to him and touched his head, and he was already cold. That broke my heart," Sugimoto said. "He was such a darling."